Specifically, it originates from the town of Villaseñor in the province of Cuenca, Spain. The name likely refers to "the lord of the manor" or "the owner of the villa".
It's important to note that the meaning of surnames can vary depending on the region and the time period in which they originated. However, the most common interpretation of Villaseñor is related to its geographical origin and the concept of ownership or authority within a village or estate.